About HMX
HMX offers a unique
approach to online learning
in medical education.
HMX courses have been envisioned and created by a highly skilled interdisciplinary team of Harvard Medical School educators and creative professionals. Together we craft curricula focused on bringing medical science concepts to life via real-world and clinical applications.
We apply the science of learning and best practices in biomedical visualization to guide the development of our courses. Our goal is to help learners focus on the understanding and application of key scientific principles, with interactive lessons and regular assessments to ensure content mastery.
Since HMX launched in 2016, we’ve delivered these unique courses to a growing number of students and professionals around the world.
What We Offer
Our courses cover key areas of medicine, from foundational science to advanced topics that are changing health care. See all available courses in our HMX Fundamentals and HMX Pro series.
Who It’s For
Learners at all levels have taken HMX courses to get ahead in their education and careers.
See what some of them have to say about the experience.
Listen In
HMX courses are designed to facilitate learning for a variety of applications. In the Learning Paths podcast, we explore educational motivations and experiences through conversations with health care students, teachers, and professionals.
Learn More
Sign up for our email list to receive information about next steps, or explore sample material in our free preview courses.
The HMX team
Michael J. Parker, MD
Associate Dean for Online Learning / Faculty Director of HMX / Assistant Professor of Medicine, Harvard Medical School
Joanne H. Muller, MA, CMI
Director of Creative Media
Sophia P. Snyder
Director of Operations, Finance, and Marketing
Jackie Kustan, MA, CMI
Senior Medical & Scientific Illustrator/Animator
Safiya Bobb
Operations Manager
Kate McDonnell-Dowling, PhD
Associate Director of Curriculum / Lecturer in Biological Chemistry and Molecular Pharmacology
Claire Stone
Senior Operations Coordinator
Matt Bunch
Principal Software Engineer
Dorothy Fatunmbi, MS, CMI
Senior Medical & Scientific Illustrator/Animator
Marysa Angelli
Operations Assistant
Andrew Pike, PhD
Immunology Curriculum Lead / Lecturer in Immunology
YeaRim Oh
Senior Curriculum Coordinator
Lauren McHale
Coordinator for Institutional and Corporate Relations
Enrique Caballero, MD
Faculty Director of Institutional and Corporate Relations
Katie Yost, MS, CMI
Senior Medical & Scientific Illustrator/Animator
Caity Anderson, PhD
Genetics Curriculum Lead / Lecturer in Genetics
Hannah Kilfoye, MS
Operations Coordinator
Harriett Gabbidon
Online Course Delivery Support Specialist
Dahye Kim
Online Course Delivery Support Specialist
Douglass McLean, MFA
Producer
Andy Twomey, MA
UI Designer
Jamie DeMotts
Educational Video Editor
Benjamin Reichman
Post Producer
Esther Ng, MS, CMI
Senior Medical & Scientific Illustrator/Animator
Faculty
Physiology
PRIMARY FACULTY
Richard M. Schwartzstein, MD
Ellen and Melvin Gordon Professor of Medicine and Medical Education, Harvard Medical School
Michael J. Parker, MD
Associate Dean for Online Learning / Faculty Director of HMX / Assistant Professor of Medicine, Harvard Medical School
CONTRIBUTING FACULTY
Melanie Hoenig, MD
Associate Professor of Medicine, Harvard Medical School / Nephrologist, Beth Israel Deaconess Medical Center
Barbara LeVarge, MD
Instructor of Medicine, Harvard Medical School / Pulmonologist, Beth Israel Deaconess Medical Center (former)
Duane Pinto, MD, MPH
Associate Professor of Medicine, Harvard Medical School / Interventional Cardiologist, Beth Israel Deaconess Medical Center
Genetics
PRIMARY FACULTY
Christine DeGennaro, PhD
Course Lead
Lecturer in Genetics, Harvard Medical School (former)
Robert Green, MD, MPH
Professor of Medicine, Harvard Medical School / Geneticist, Brigham and Women’s Hospital
Carrie Blout, MS, CGC, LGC
Senior Genetic Counselor, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
Calum MacRae, MD, PhD
Professor of Medicine, Harvard Medical School / Cardiologist, Brigham and Women’s Hospital
Jonathan Picker, MD, PhD
Assistant Professor of Medicine, Harvard Medical School / Medical Geneticist, Boston Children’s Hospital
Huma Rana, MD
Assistant Professor of Medicine, Harvard Medical School / Medical Geneticist, Dana-Farber Cancer Institute
Immunology
PRIMARY FACULTY
Andrew Lichtman, MD, PhD
Professor of Pathology, Harvard Medical School
Shiv Pillai, MD, PhD
Professor of Medicine and Health Sciences and Technology, Harvard Medical School
Marshall Thomas, PhD
Lecturer in Genetics, Harvard Medical School (former)
CONTRIBUTING FACULTY
Jules Dienstag, MD
Carl W. Walter Professor of Medicine, Harvard Medical School / Gastroenterologist, Massachusetts General Hospital
Keith Flaherty, MD
Professor of Medicine, Harvard Medical School / Hematologist/ Oncologist, Massachusetts General Hospital
Joseph Merola, MD, MMSc
Associate Professor, Harvard Medical School / Dermatologist, Brigham and Women’s Hospital
David Sloane, MD
Instructor, Harvard Medical School / Allergist and Immunologist, Brigham and Women’s Hospital
Biochemistry
PRIMARY FACULTY
Kevin Bonham, PhD
Course Lead
Lecturer in Microbiology and Immunobiology, Harvard Medical School (former)
Ole-Petter (“OP”) Hamnvik, MB BCh BAO, MMSc
Assistant Professor of Medicine, Harvard Medical School / Endocrinologist, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
Marie E. McDonnell, MD
Lecturer on Medicine, Harvard Medical School / Endocrinologist, Brigham and Women’s Hospital
Eli Miloslavsky, MD
Assistant Professor of Medicine, Harvard Medical School / Rheumatologist, Massachusetts General Hospital
Pharmacology
PRIMARY FACULTY
Kate McDonnell-Dowling, PhD
Course Lead
Lecturer in Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Associate Director of Curriculum, HMX
James P. Rathmell, MD
Primary Clinical Faculty
Leroy D. Vandam Professor of Anesthesia, Harvard Medical School / Chair of the Department of Anesthesiology, Perioperative and Pain Medicine at Brigham and Women’s Hospital
David E. Golan, MD, PhD
Primary Advising Faculty
Professor of Biological Chemistry and Molecular Pharmacology, Harvard Medical School /
Dean for Research Operations and Global Programs, Harvard Medical School /
George R. Minot Professor of Medicine, Department of Medicine, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
Nikhil C. Munshi, MD
Professor of Medicine, Harvard Medical School / Medical Oncologist, Dana-Farber Cancer Institute
Pritesh Topiwala, MD
Instructor in Anesthesia, Brigham and Women’s Hospital / Harvard Medical School
Cancer Genomics and Precision Oncology
PRIMARY FACULTY
Christine DeGennaro, PhD
Course Lead
Lecturer in Genetics, Harvard Medical School (former)
Marios Giannakis, MD, PhD
Assistant Professor of Medicine, Harvard Medical School / Medical Oncologist, Dana-Farber Cancer Institute
Robert Green, MD, MPH
Professor of Medicine, Harvard Medical School / Geneticist, Brigham and Women’s Hospital
Carrie Blout, MS, CGC, LGC
Senior Genetic Counselor, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
George Demetri, MD
Professor of Medicine, Harvard Medical School / Medical Oncologist, Dana-Farber Cancer Institute
William Hahn, MD, PhD
William Rosenberg Professor of Medicine, Harvard Medical School and Dana-Farber Cancer Institute / Medical Oncologist, Dana-Farber Cancer Institute
William Kaelin, MD
Sidney Farber Professor of Medicine, Harvard Medical School and Dana-Farber Cancer Institute / Nobel Laureate in Physiology or Medicine
Tali Mazor, PhD
Computational Biologist, Dana-Farber Cancer Institute
Matthew Meyerson, MD, PhD
Professor of Pathology, Harvard Medical School and Dana-Farber Cancer Institute / Medical Oncologist, Dana-Farber Cancer Institute
Jonathan Nowak, MD, PhD
Assistant Professor of Pathology, Brigham and Women’s Hospital / Molecular Pathologist, Brigham and Women’s Hospital
Huma Rana, MD
Assistant Professor of Medicine, Harvard Medical School / Medical Geneticist, Dana-Farber Cancer Institute
Alice Shaw, MD, PhD
Professor of Medicine, Harvard Medical School / Thoracic Oncologist, Massachusetts General Hospital
Lynette Sholl, MD
Associate Professor of Pathology, Harvard Medical School / Pathologist, Brigham and Women’s Hospital
Immuno-oncology
PRIMARY FACULTY
Andrew Lichtman, MD, PhD
Professor of Pathology, Harvard Medical School
Shiv Pillai, MD, PhD
Professor of Medicine and Health Sciences and Technology, Harvard Medical School
Andrew Pike, PhD
Lecturer in Immunology, Harvard Medical School / Curriculum Fellow in Online Learning
CONTRIBUTING FACULTY
Keith Flaherty, MD
Professor of Medicine, Harvard Medical School / Hematologist/ Oncologist, Massachusetts General Hospital
Gordon Freeman, PhD
Professor of Medicine, Harvard Medical School
Timothy Graubert, MD
Professor of Medicine, Harvard Medical School / Hematologist/Oncologist, Massachusetts General Hospital
F. Stephen Hodi, MD
Professor of Medicine, Harvard Medical School / Medical Oncologist, Dana-Farber Cancer Institute
Marcela Maus, MD, PhD
Assistant Professor of Medicine, Harvard Medical School / Hematologist/Oncologist, Massachusetts General Hospital
Nikhil C. Munshi, MD
Professor of Medicine, Harvard Medical School / Medical Oncologist, Dana-Farber Cancer Institute
Lynette Sholl, MD
Associate Professor of Pathology, Harvard Medical School / Pathologist, Brigham and Women’s Hospital
Scott Rodig, MD, PhD
Associate Professor of Pathology, Harvard Medical School / Pathologist, Dana-Farber Cancer Institute
Genetics – Essentials
PRIMARY FACULTY
Christine DeGennaro, PhD
Course Lead
Lecturer in Genetics, Harvard Medical School (former)
Robert Green, MD, MPH
Professor of Medicine, Harvard Medical School / Geneticist, Brigham and Women’s Hospital
Carrie Blout, MS, CGC, LGC
Senior Genetic Counselor, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
George Church, PhD
Robert Winthrop Professor of Genetics, Harvard Medical School / Professor of Health Sciences and Technology, Harvard Medical School
Isaac (“Zak”) Kohane, MD, PhD
Marion V. Nelson Professor of Biomedical Informatics, Harvard Medical School / Lawrence J. Henderson Professor of Pediatrics and Health Sciences and Technology, Harvard Medical School
Calum MacRae, MD, PhD
Professor of Medicine, Harvard Medical School / Cardiologist, Brigham and Women’s Hospital
Heidi Rehm, PhD
Professor of Pathology, Harvard Medical School / Chief Genomics Officer, Massachusetts General Hospital
Pharmacology – Essentials
PRIMARY FACULTY
Kate McDonnell-Dowling, PhD
Course Lead
Lecturer in Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Associate Director of Curriculum, HMX
James P. Rathmell, MD
Primary Clinical Faculty
Leroy D. Vandam Professor of Anesthesia, Harvard Medical School / Chair of the Department of Anesthesiology, Perioperative and Pain Medicine at Brigham and Women’s Hospital
David E. Golan, MD, PhD
Primary Advising Faculty
Professor of Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Dean for Research Operations and Global Programs, Harvard Medical School /
George R. Minot Professor of Medicine, Department of Medicine, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
Elizabeth Garner, MD, MPH
Chief Medical Officer, ObsEva SA / Harvard Medical School alumna
Nikhil C. Munshi, MD
Professor of Medicine, Harvard Medical School / Medical Oncologist, Dana-Farber Cancer Institute
Vishal Patel, PhD
Chief of Staff, Novartis Institutes for BioMedical Research / Harvard Medical School alumnus
Peter Sorger, PhD
Otto Krayer Professor of Systems Pharmacology, Harvard Medical School
Pritesh Topiwala, MD
Instructor in Anesthesia, Brigham and Women’s Hospital / Harvard Medical School
Novel Therapies for Chronic Inflammation, Autoimmunity, and Allergy
PRIMARY FACULTY
Andrew Lichtman, MD, PhD
Professor of Pathology, Harvard Medical School
Shiv Pillai, MD, PhD
Professor of Medicine and Health Sciences and Technology, Harvard Medical School
Andrew Pike, PhD
Lecturer in Immunology, Harvard Medical School / Curriculum Fellow in Online Learning
CONTRIBUTING FACULTY
Michael Dougan, MD, PhD
Assistant Professor of Medicine, Harvard Medical School / Gastroenterologist, Massachusetts General Hospital
Aidan Long, MD
Associate Professor of Medicine, Harvard Medical School / Allergist, Massachusetts General Hospital
Joseph Merola, MD, MMSc
Associate Professor, Harvard Medical School / Dermatologist, Brigham and Women’s Hospital
Deepak Rao, MD, PhD
Assistant Professor of Medicine, Harvard Medical School / Rheumatologist, Brigham and Women’s Hospital
Michael Weinblatt, MD
John R. Riedman Professor of Medicine, Harvard Medical School / Rheumatologist, Brigham and Women’s Hospital
Genetic Testing and Sequencing Technologies
PRIMARY FACULTY
Caity Anderson, PhD
Course Lead
Lecturer in Genetics, Harvard Medical School / Curriculum Fellow in Online Learning
David Miller, MD, PhD
Primary Clinical Faculty
Associate Professor of Pediatrics, Harvard Medical School / Medical Geneticist, Boston Children’s Hospital
CONTRIBUTING FACULTY
Mark Daly, PhD
Associate Professor of Medicine, Harvard Medical School / Geneticist, Massachusetts General Hospital
Christine DeGennaro, PhD
Lecturer in Genetics, Harvard Medical School (former)
Amit Khera, MD, MSc
Assistant Professor of Medicine, Harvard Medical School / Cardiologist, Massachusetts General Hospital
Cynthia Morton, PhD
Professor of Pathology, Harvard Medical School / Medical Geneticist, Brigham and Women’s Hospital
Anne O’Donnell Luria, MD, PhD
Instructor in Pediatrics, Harvard Medical School / Medical Geneticist, Boston Children’s Hospital
Michelle Pacione, EdM, MS, LCGC
Genetic Counselor, Brigham and Women’s Hospital
Timothy Yu, MD, PhD
Associate Professor of Pediatrics, Harvard Medical School / Neurologist, Boston Children’s Hospital
Drug Delivery
PRIMARY FACULTY
Kate McDonnell-Dowling, PhD
Course Lead
Lecturer in Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Associate Director of Curriculum, HMX
Daniel Kohane, MD, PhD
Primary Faculty
Professor of Anesthesia, Harvard Medical School / Pediatric Critical Care Physician, Boston Children’s Hospital
Philip Cole, MD, PhD
Primary Advising Faculty
Professor of Medicine and Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Senior Investigator, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
Michael Agus, MD
Associate Professor of Pediatrics, Harvard Medical School / Pediatric Endocrinologist and Pediatric Critical Care Physician, Boston Children’s Hospital
Katharine Garvey, MD, MPH
Assistant Professor of Pediatrics, Harvard Medical School / Pediatric Endocrinologist, Boston Children’s Hospital
Ana Jaklenec, PhD
Research Scientist and Principal Investigator, MIT
Jeffrey Karp, PhD
Professor of Medicine, Harvard Medical School, Brigham and Women’s Hospital
Robert Langer, ScD
David H. Koch Institute Professor, MIT / Faculty, Harvard-MIT Division of Health Sciences and Technology
Xueguang Lu, PhD
Postdoctoral Research Fellow, MIT
Morteza Sarmadi, MSc
PhD candidate, MIT
Garry Steil, PhD
Assistant Professor of Pediatrics, Harvard Medical School / Physiologist and Research Engineer, Boston Children’s Hospital
C. Giovanni Traverso, MB, BChir, PhD
Assistant Professor of Medicine, Harvard Medical School / Gastroenterologist, Brigham and Women’s Hospital
ADDITIONAL CONTRIBUTORS
Yang Li, PhD
Research Fellow, Boston Children’s Hospital
Wei Zhang, PhD
Instructor in Anesthesiology, Boston Children’s Hospital
Vaccines and Viral Immunology
PRIMARY FACULTY
Andrew Lichtman, MD, PhD
Professor of Pathology, Harvard Medical School
Shiv Pillai, MD, PhD
Professor of Medicine and Health Sciences and Technology, Harvard Medical School
Andrew Pike, PhD
Lecturer in Immunology, Harvard Medical School / Curriculum Fellow in Online Learning
CONTRIBUTING FACULTY
Galit Alter, PhD
Professor of Medicine, Harvard Medical School
Dan Barouch, MD, PhD
William Bosworth Castle Professor of Medicine, Harvard Medical School
Jules Dienstag, MD
Carl W. Walter Professor of Medicine, Harvard Medical School / Gastroenterologist, Massachusetts General Hospital
Stephen C. Harrison, PhD
Giovanni Armenise-Harvard Professor of Basic Medical Sciences, Harvard Medical School
Kathryn E. Stephenson, MD, MPH
Assistant Professor of Medicine, Harvard Medical School
Bruce Walker, MD
Phillip T. and Susan M. Ragon Professor of Medicine, Harvard Medical School
Drug Discovery and Development
PRIMARY FACULTY
Kate McDonnell-Dowling, PhD
Lecturer in Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Associate Director of Curriculum, HMX
Mark Goldberg, MD
Lecturer in Medicine, Harvard Medical School / Attending Physician in Hematology, Brigham and Women’s Hospital / Member of Board of Directors at Avacta Group PLC
Mark Namchuk, PhD
Senior Lecturer in Biological Chemistry and Molecular Pharmacology, Harvard Medical School / Executive Director of Therapeutics Translation in Department of Biological Chemistry and Molecular Pharmacology, Harvard Medical School
William Clarke, MD
Professor of Anesthesia, Harvard Medical School / Pediatric Critical Care Physician, Boston Children’s Hospital
Paul Ridker, MD, MPH
Eugene Braunwald Professor of Medicine, Harvard Medical School / Senior Physician, Brigham and Women’s Hospital / Director of the Center for Cardiovascular Disease Prevention, Brigham and Women’s Hospital
CONTRIBUTING FACULTY
David Altshuler, MD, PhD
Executive Vice President of Global Research and Chief Scientific Officer, Vertex Pharmaceuticals / Senior Lecturer in Genetics, Harvard Medical School
Julie Buring, DSc
Professor of Medicine, Harvard Medical School / Epidemiologist, Brigham and Women’s Hospital
Rory Collins, FRS, FMedSci
Professor of Medicine and Epidemiology, University of Oxford / Principal Investigator and Chief Executive, UK Biobank
Elizabeth Garner, MD, MPH
Chief Medical Officer and Senior Vice President, Agile Therapeutics / Owner and President, KNI Health Consultants / Harvard Medical School alumna
Sam Moskowitz, MD
Executive Medical Director, Vertex Pharmaceuticals / Lecturer, Harvard Medical School
Paul Negulescu, PhD
Senior Vice President and Site Head of San Diego Research, Vertex Pharmaceuticals
Saurabh Saha, MD, PhD
Chief Executive Officer, Centessa Pharmaceuticals / Associate Member, Harvard Medical School
Priya Singhal, MD, MPH
Senior Vice President and Head of Global Safety & Regulatory Sciences, Biogen / Harvard Medical School alumna
Frank Slack, PhD
Shields Warren Mallinckrodt Professor, Harvard Medical School / Director of the Institute for RNA Medicine, Beth Israel Deaconess Medical Center
Vishal Vaidya, PhD
Senior Director and Head of Clinical Biomarker Technologies, Pfizer / Associate Professor, Harvard Medical School
ADDITIONAL CONTRIBUTORS
Juli Coraor Fried
MD/PhD candidate, Harvard Medical School
Michael Poeschla
MD/PhD candidate, Harvard Medical School
Gene Therapy
PRIMARY FACULTY
Caity Anderson, PhD
Lecturer in Genetics, Harvard Medical School / Genetics Curriculum Lead, HMX
Connie Cepko, PhD
Bullard Professor of Genetics and Neuroscience, Harvard Medical School / Howard Hughes Medical Institute Investigator
Christine Duncan, MD
Associate Professor of Pediatrics, Harvard Medical School / Senior Physician, Dana-Farber/Boston Children’s Cancer and Blood Disorders Center / Medical Director of Clinical Research and Clinical Development, Gene Therapy Program, Boston Children’s Hospital
CONTRIBUTING FACULTY
Jason Comander, MD, PhD
Assistant Professor of Ophthalmology, Harvard Medical School / Director of the Inherited Retinal Disorders Service, Massachusetts Eye and Ear
David Corey, PhD
Bertarelli Professor of Translational Medical Science, Harvard Medical School
Florian Eichler, MD
Associate Professor of Neurology, Harvard Medical School / Neurologist and Director of the Leukodystrophy Service, Massachusetts General Hospital
Erica Esrick, MD
Instructor in Pediatrics, Harvard Medical School / Pediatric Hematologist, Boston Children’s Hospital
Rachel Huckfeldt, MD, PhD
Assistant Professor of Ophthalmology, Harvard Medical School / Ophthalmologist, Massachusetts Eye and Ear / Director of the Inherited Retinal Degenerations Fellowship, Massachusetts Eye and Ear
Patricia Musolino, MD, PhD
Assistant Professor of Neurology, Harvard Medical School / Neurologist, Massachusetts General Hospital / Co-Director of the Pediatric Stroke and Cerebrovascular Service, Massachusetts General Hospital / Faculty, Center for Genomic Medicine, Massachusetts General Hospital
David Williams, MD
Leland Fikes Professor of Pediatrics, Harvard Medical School / Chief of the Division of Hematology/Oncology, Boston Children’s Hospital / Associate Chairman, Department of Pediatric Oncology, Dana-Farber Cancer Institute
ADDITIONAL CONTRIBUTORS
Jillian F. Wise, PhD
Instructor in Investigation, Massachusetts General Hospital / Harvard Medical School / Broad Institute
Jordan Said, MD
Internal Medicine Resident, Brigham and Women’s Hospital / Harvard Medical School
Allie Stanton
PhD Candidate, Harvard Medical School
HMX Moderators
Elliot Akama-Garren
Harvard Medical School student
MD/PhD Candidate, 2025
Adam Berger
Harvard Medical School student
MD/PhD Candidate, 2026
Michael Chang, MD
PGY-1
Brigham and Women’s Hospital
Juli Coraor Fried
Harvard Medical School student
MD/PhD Candidate, 2024
Carlos Donado
Postdoctoral Research Fellow
Brigham and Women’s Hospital / Harvard Medical School
Hannah Hoffman
Harvard Medical School student
MD/PhD Candidate, 2029
Ameya Kirtane
Instructor in Medicine
Brigham and Women’s Hospital / Harvard Medical School
Gha Young Lee
Harvard Medical School student
MD Candidate, 2024
Yang Li
Postdoctoral Research Fellow
Boston Children’s Hospital / Harvard Medical School
Paris Pallis
Immunology,
Harvard Medical School
PhD Candidate
Michael Poeschla
Harvard Medical School student
MD/PhD candidate, 2027
Michael Quezada
Harvard Medical School student
MD/PhD Candidate, 2029
Jacob Ronald Anderson
Harvard Medical School student
MD/PhD Candidate, 2026
Justine Rutter
Harvard Medical School student
MD/PhD Candidate, 2028
Jordan Said, MD
Internal Medicine Resident
Brigham and Women’s Hospital / Harvard Medical School
Priya Shah, MD
Pediatrics Resident
Children’s Hospital of Philadelphia
Harvard Medical School Alumna
Blake Smith
Harvard Medical School student
MD/PhD Candidate, 2026
Jillian F. Wise, PhD
Postdoctoral Research Fellow and Visiting Scholar
University of Oslo / Massachusetts General Hospital / Harvard Medical School / Broad Institute
